Athermal quasistatic simple shear of the 2D Voronoi tissue model under Lees–Edwards boundaries. With K_A = 0 the energy is perimeter-only: E = Σ K_P (P_i − P_0)². Default box side L = √N so ⟨A⟩ = A₀ = 1.
Each strain step applies an affine shear x → x + Δγ·y, wraps under LEBC, then FIRE-minimises to mechanical equilibrium. Shear stress σ_xy is measured from the virial/analytical form (1/L²) Σ_e T_e dx_e dy_e / |e|, which does not spike at T1s. T1 events are detected by changes in the Delaunay edge set; the four cells of each T1 quartet flash red briefly.
Huang, J., Cochran, J., Fielding, S., Marchetti, M., & Bi, D. Shear-Driven Solidification and Nonlinear Elasticity in Epithelial Tissues. Phys. Rev. Lett. 128, 178001 (2022).
Nguyen, A. Q., Huang, J., & Bi, D. Origin of yield stress and mechanical plasticity in model biological tissues. Nat. Commun. 16, 3260 (2025).